Bachelor of Medicine, Edinburgh University, 1964. Doctor of Philosophy, Edinburgh University, 1976.
Lecturer in therapeutics Edinburgh University, United Kingdom, 1968-1972. Senior research fellow Leiden University, United Kingdom, 1972-1973. Senior lecturer Edinburgh University, United Kingdom, 1973-1990.
Professor medical genetics Nottingham University, United Kingdom, since 1990. Genetic advisor Association British Insurers, since 1996. Member Government Genetics and Insurance Committee, since 1999.
Major Royal Army Medical Corps, 1966-1985. Fellow Royal College Physicians. Member Scottish Down Syndrome Associations (past chair), Cystic Fibrosis Research Trust (past chair).
Married Patricia Hawco, December 8, 1967 (divorced April 1980). Children: Elspeth, Alison, Hugh. Married Arlene Rose Conway, May 17, 1980.