HND, Manchester Polytechnic, 1971. MIBiol, Ewell College, London, 1973. Doctor of Philosophy, Aston University, Birmingham, England, 1976.
Research associate Ministry of Defense, Wiltshire, England, 1970—1971, Ministry of Agriculture, Windsor, 1971—1973. Postdoctoral fellow University of California at Los Angeles, 1976—1979, Max Planck Institute, Tubingen, Germany, 1979—1980, Stanford University, 1980—1981. Assistant professor biology University of California at Los Angeles, since 1981.
Member of American Society Microbiology, Biophysics Society, Institute Biology.
Married Bernadine Wisnieski, May 28, 1980.