Bachelor of Arts Zoology, Cornell Univercity, 1956; Doctor of Philosophy, Yale University, 1960.
Assistant professor zoology, U. Iowa, Iowa City, 1962-1965; associate professor, U. Iowa, Iowa City, 1965-1971; professor biology, U. Iowa, Iowa City, since 1971.
Member American Association for the Advancement of Science, Society Development Biologists, Society Protozoologists.
Married Anne Wijtske Koopmans, December 21, 1961. Children– Rebecca Eva, Martin David.