Background
He was born in Brechin in 1928 and was educated at Brechin High School and the University of Edinburgh, where he read medicine.
He was born in Brechin in 1928 and was educated at Brechin High School and the University of Edinburgh, where he read medicine.
University of Edinburgh.
After a spell as a physician he returned to university to obtain a Doctor of Philosophy then took lectureships at the University of Edinburgh and the University of Glasgow. In 1969 he was appointed to the Chandos Chair of Physiology at the University of Street Andrews and held the post for 24 years until 1993. From 1998 until 2003 he led the Rowett Research Institute at the University of Aberdeen.
He was elected as a Fellow of the Royal Society of Edinburgh in 1986.