Background
Varner, Joseph Elmer was born on October 7, 1921 in Nashport, Ohio, United States. Son of George Ezra and Inez Charlotte (Gladden) Varner.
Varner, Joseph Elmer was born on October 7, 1921 in Nashport, Ohio, United States. Son of George Ezra and Inez Charlotte (Gladden) Varner.
Bachelor of Science, Ohio State University, 1942. Master of Science, Ohio State University, 1943. Doctor of Philosophy, Ohio State University, 1949.
Docteur Honoris Causa (honorary), Nancy University, France, 1977.
Chemist, Owens Corning, Newark, 1943-1944; analytical chemist, Battelle Memorial Institute, Columbus, Ohio., 1946-1947; from assistant professor to professor biology, Ohio State University, Columbus, Ohio., 1950-1961; postdoctoral, California Institute Technology, Pasadena, 1953-1954; National Science Foundation fellow, Cambridge U., England, 1959-1960; research scientist, Martin-Marietta, Baltimore, 1961-1965; professor biochemistry, Michigan State University, East Lansing, 1965-1973; National Science Foundation fellow, U. Washington, Seattle, 1971-1972; professor biology, Washington University, St. Louis, 1973-1995; American Cancer Society scholar, Washington University, St. Louis, 1980-1981.
Fellow American Association for the Advancement of Science. Member National Academy of Sciences, American Society Plant Physiologists (president 1969, Stephen Hales Prize 1990), Society Developmental Biology (president 1986), American Academy Arts and Science.
Married Carol Roberta Dewey, June, 1945 (divorced 1971). Children— Lee, Lynn, Karen, Beth. Married Jane Elanor Burton, June, 1976.