Bachelor, Yale University, 1966; Master of Science, Lehigh University, 1976; Doctor of Philosophy, Cambridge (England) University, 1983.
Volunteer, Peace Corps, Western Caroline Islands, 1966-1969; production manager, Pennsylvania Pacific Corporation, Warminster, 1970-1972; research assistant department biology, Lehigh University, Bethlehem, Pennsylvania, 1975-1976; scientist department zoology, U. Oxford, England, 1983-1987; scientist, Fraunhofer Institute, Hannover, Germany, 1987-1991; scientist, Department of Agriculture, Hannover, Germany, since 1991.
Member American Association for the Advancement of Science, New York Academy Science.
Married Susan Caspari, May 16, 1970. Children: Joseph C., James W., John D.