Bachelor, Yale University, 1970; Master of Arts, Doctor of Philosophy, Harvard University, 1976.
With United States Congress, Office of Technology Assessment, 1976—1977, University California, San Francisco. Assistant professor, department physiology Washington University Medical School, St. Louis, 1980-1985, associate professor, 1985-1989, alumni endowed professor neurobiology, department anatomy and neurobiology, 1989. Professor molecular and cellular biology Harvard University, Cambridge, Massachusetts, Paul J. Finnegan Family director, Center for Brain Neuroscience, Department Molecular and Cellular Biology, since 2005.
Member neurology study section National Institutes of Health, Washington, 1988. Member national advisory council, National Institute Neurological Disorders and Stroke, National Institutes of Health, 1999-2003. Member advisory committee, human embryonic stem cell research, National Research Council and Institute Medicine, since 2006.
Member scientific advisory board Stowers Institute for Medical Research, since 2006, Searle Scholar program, Max-Planck Institute for Neurobiology in Munich, Howard Hughes Medical Institute.
Fellow American Association for the Advancement of Science, American Academy Arts & Sciences. Member National Institutes of Health (board science counselor since 1993), Society for Neurosci. (councilor since 1990), Muscular Dystrophy Association (scientific advisory board since 1991), National Academy of Sciences.
Married Susan Corcoran, December 27, 1982. Children: Jesse, Amelia.