Education
He studied medicine at the Universities of Würzburg and Freiburg, earning his medical doctorate in 1881.
He studied medicine at the Universities of Würzburg and Freiburg, earning his medical doctorate in 1881.
Following graduation, he furthered his education at Tübingen with pathologist Ernst Ziegler and in Vienna with dermatologists Moriz Kaposi and Isidor Neumann. From 1885 onward, he was a practicing dermatologist in Wiesbaden. As a botanist he specialized in studies of the genus Hieracium.
He worked closely with botanist August Schlickum (1867–1946), with whom he conducted scientific excursions in the Rhineland and the AllgäUniversity
His herbarium of around 20,000 items was acquired in 1929 and 1935 by the botanical museum at Berlin-Dahlem. "Touton giant cell": A xanthoma cell in which the multiple nuclei are grouped around a small island of non-foamy cytoplasm.