Bachelor of Science, University Helsinki, 1963. Master of Public Health, University Helsinki, 1965. Doctor of Philosophy, University Helsinki, 1973.
Researcher University Helsinki, 1966-1983, assistant teacher department zoology, 1980-1992, senior curator Finnish Museum Natural History, 1981-1982, 93—. Researcher Biosystematics Institute, Ottawa, Canada, 1985-1986, 92-93. Specialist on invertebrates Finnish Broadcasting Company, since 1974.
Vice president Lepidopterists Society, New Haven, 1994-1996. Member council Societas Europaea Lepidopterologica, Basel, Switzerland, 1986-1992, vice president, 1990-1992. Fellow Royal Entomological society.
Member Russian Entomological Society (honorary), Polish Entomological Society (honorary), Finnish Lepidopterol. Society (honorary, chairman 1980-1984, professor honoris causa).
Married Tarja Liisa Lahtinen, 1966 (divorced 1980). Children: Melina, Matti. Married Maarit Irmeli Louekari, December 23, 1983 (divorced 2000).
1 child, Reija.