Doctor of Medicine, University Tokyo, 1956. Doctor Medical Science, University Tokyo, 1961. Master of Arts (honorary), Yale University, 1983.
Assistant professor medicine Harvard Medical School, Boston, 1969-1973. Senior research scientist Yale University, New Haven, 1973-1982, professor genetics, 1983-1994, professor emeritus, since 1995. Member biochemistry study section National Institutes of Health, Bethesda, Maryland, 1983-1984.
Member American Society Biological Chemistry, American Society Human Genetics, Society Inborn Metabolic Disorders.
Married Tomoko Hasegawa, November 5, 1954. Children: Atau, Elly Margaret.