Doctor of Philosophy, Uppsala University, Sweden, 1971. Doctor of Medicine, Uppsala University, Sweden, 1972.
Associate professor pathology Uppsala University, 1971-1976. Senior research fellow Swedish Cancer Society, Uppsala, 1976-1980. Professor cellular pathology Uppsala University, since 1980.
Consultant Sandoz, Vienna, 1981-1985, Pharmacia, Uppsala, 1981-1993.
Chairman, Swedish Cancer Society, since 1992. Member American Association Cancer Research, Norwegian Academy of Sciences.
Married Karin Edsjo, May 22, 1965. Children: Lisa, Anders.