Bachelor, Lawrence University, 1973; Doctor of Philosophy, University of California Santa Cruz, 1977.
Assistant professor department biochemistry, McGill University, Montreal, Quebec, 1978-1981; assistant professor departments biochemistry & molecular biology, U. Florida, Gainesville, 1981-1985; associate professor neuroscience, U. Florida, Gainesville, 1985-1986; associate professor department molecular physiology, Baylor College Medicine, Houston, 1986-1991; professor department cell biology, Baylor College Medicine, Houston, 1991-1995; professor, chair animal cell biology department, Durham (England) University, since 1995. Past member National Institutes of Health study sections biology, neurology, Bethesda, Maryland., science advising committee National Multiple Sclerosis Society.
Children: Alexei, Nicholas. Married L. Karaviti, September 21, 1991. Children: Philip, Eleni.