Education
Staatsexamen, University Darmstadt, Germany, 1972. Doctor rerum naturalium (Doctor in Sciences), University Darmstadt, Germany, 1975. Doctor rerum naturalium habilitatus, Universität Würzburg, Germany, 1983.
Staatsexamen, University Darmstadt, Germany, 1972. Doctor rerum naturalium (Doctor in Sciences), University Darmstadt, Germany, 1975. Doctor rerum naturalium habilitatus, Universität Würzburg, Germany, 1983.
Wissenschaften angestellter University Darmstadt, Botany Institute, 1975—1977. Postdoctoral fellow Australian National University, RSBS, Canberra, 1977—1979. Postdoctoral research associate University Wisconsin, Madison, 1980.
Habilitation fellow University Wurzburg, 1981—1983. Heisenberg fellow University Wurzburg, Botany Institute, 1983—1985, professor, 1985—1991. Staff scientist Smithsonian Tropical Research Institute, Panama City, Panama, 1991—1998, senior staff member Panama, since 1998.