Bachelor, Kyoto University, 1965. Master of Agriculture, Kyoto University, 1967. Master of Arts, State University of New York, Stony Brook, 1971.
Doctor of Philosophy, University Kansas, 1972.
Research fellow University British Columbia, Vancouver, Canada, 1971-1973. Assistant professor Purdue University, West Lafayette, Indiana, 1973-1978. Associate professor University Tsukuba, Ibaraki, Japan, 1978-1988, professor Japan, 1988—2006, emeritus professor Japan, since 2006, chair graduate program biology Japan, 1994-1997, provost graduate program Japan, 2003—2006.
Member Ecolog. Society American, Society Population Ecology (president 1997), Japanese Society Ecology, Sigma Xi.
Married Isuzu Sano, January 28, 1969. 1 child, Rika.