Bachelor, Albion College, 1957. Master of Science, University Chicago, 1962. Doctor of Philosophy, University California, Santa Barbara, 1967.
Assistant professor, associate professor biology University Puerto Rico, Rio Piedras, 1967-1976. Associate professor biology and marine science Long Island University, Southampton, New York, 1976, professor, since 1978.
Fellow American Association for the Advancement of Science. Member Phycological Society of America (treasurer l985-88, trustee l989—), International Phycological Society, American Institute Biological Sciences (representative to council).