Career
After education at Leighton Park School and Cambridge University he became an academic at Cambridge University. He was an early Mendelian geneticist who discovered sex linkage, while writing up the results of the Reverend G.H. Raynor on the magpie moth Abraxas grossulariata. He later wrote a number of books on Mendelian genetics and on sex determination.
He was appointed assistant to the Superintendent of the Cambridge University Museum of Zoology in June 1902, and himself filled this position from 1909 to 1914.
He died of sarcoma in 1920, and William Bateson wrote his obituary in Nature. His book, is notable for explicitly dismissing Lamarckian inheritance.