Bachelor of Science, W Texas State University, 1961. Master of Science, University Arkansas, 1963. Doctor of Philosophy, University Louisville, 1966.
Assistant professor to associate professor zoology, University Oklahoma, 1966-1975; assistant director biological studies, University Oklahoma, 1967-1969; director biological station, University Oklahoma, 1969-1974; professor zoology and director biology, University Oklahoma, since 1969; department chairman zoology, University Oklahoma, 1980-1984. Instructor zoology Catherine Spalding College, 1965-1966. President Southwestern Association naturalists, 1977-1979.
Member American Fisheries Society, American Society Ichthyologists & Herpetologists.
Married, 1983; children.