Bachelor of Science, St. John's University, 1957. Master of Science, Creighton University, 1959. Doctor of Philosophy, University Missouri, 1964.
Postdoctorate, Mayo Clinic, Rochester, Minnesota, 1964-1966; postdoctorate, St. Jude Children's Hospital, Memphis, 1966-1968; assistant member, St. Jude Children's Hospital, Memphis, 1968-1970; assistant professor microbiology, Bowman Gray School Medicine, Winston Salem, North Carolina, 1970-1974; associate professor, Bowman Gray School Medicine, Winston Salem, North Carolina, 1974-1980; professor, Bowman Gray School Medicine, Winston Salem, North Carolina, since 1980. Research consultant National Institutes of Health, Bethesda, Maryland., since 1980, North Carolina Association for biomedical Research, Research Triangle Park, since 1985.
With United States Army, 1959-1960. Member American Society for Microbiology, American Society for Virology, International Society Antiviral Research, International Society for Acuired Immune Deficiency Syndrome Research.
Married JoAnn Dorothy Martinson, August 22, 1959. Children: Gregory, Gary, Stephen, Scott.