Bachelor of Science in Biochemistry, University Complutense, Madrid, 1981. Doctor of Philosophy, University Autonoma, Madrid, 1985.
Postdoctoral staff, C.B.M. (Spanish National Research Council-Universidad Autónoma Metropolitana), Madrid, 1985-1987; postdoctoral staff, M.R.C., Cambridge, England, 1987-1991; group leader, European Molecular Biology Laboratory, Heidelberg, Germany, since 1992.
Second lieutenant Spanish Army (infantry), 1982-1983. Member New York Academy of Sciences.
Married Isabel Vernos, June 29, 1985. Children: Miguel, Laura.