Laurea in Biological Sciences, University Messina, Italy, 1986. Doctor of Philosophy in Environmental Marine Sciences, University Genoa, Italy, 1992.
Visiting scientist department plant biology and ecology University St. Andrews, England, 1986-1987. Trainee, biologist department organic and biological chemistry University Messina, Italy, 1987-1988. Visiting scientist department botany Photosynthesis Research Laboratory University Florida, Gainesville, 1988-1991, adjunct assistant, since 1991.
Postdoctoral fellow Robert Hill Institute, University Sheffield, England, 1991-1992. Visiting scientist department ecology and evolution biology Monash University, Melbourne, Australia, 1996, 97. Researcher faculty science University Ancona, Italy, since 1994.
Visiting scientist IMCS, Rutgers University, New Jersey, 1999-2000.
Member American Society Plant Physiologists (travel award 1997), International Phycological Society, Phycological Society of America.