Bachelor of Arts in Biophysics, Amherst College, 1964. Doctor of Philosophy in Biophysics, Johns Hopkins University, 1969.
Research fellow Australian National University, Canberra, 1969-1971. Assistant professor department molecular, cellular and development biology University Colorado, Boulder, 1971-1977, associate professor, 1977-1982, professor, 1982—2008, chairman department, 1983—1987, emeritus professor, since 2008, associate vice chancellor for academy affairs, 1988—1997, chief information officer, 1996—1997, faculty fellow information technical services, 1997-1998, director academy development BP Center for Visualization, 2003—2005. Scientific consultant Wills Foundation, 1981-1991.
Consultant, member board science advisors Columbine Venture Fund, Denver, 1984-1994, Photometrics, Tucson, 1987-1989. Member academy advisory board higher education Apple Computing, 1997-1998. Co-founder 3D Embodiment, LLC, managing partner since 2005.
Board of directors Congregation Har Ha-Shem, Boulder, 1976-1980, president, 1978, 79, Cmty. Access television of Boulder, 1996-1997. Member American Association for the Advancement of Science, AAHE, American Association of University Professors, Association Research in Vision and Ophthalmology (secretary chairman 1981), Society Neurosci., Sigma Xi.
Married Alma Hermine Heller, June 27, 1964. Children: Lila Rachel, Miriam Rebecca.