Bachelor of Arts Chicago, 1963. Doctor of Philosophy, University Chicago, 1968.
National Science Foundation postdoctoral fellow Yale University, 1968-1970. Assistant professor biology University Rochester (New York ), 1970-1975, associate professor, 1975-1980, professor, since 1980, chairman department biology, since 1981. Member biological development panel National Science Foundation, American Cancer Society.
Member American Society Cell Biology, American Association for the Advancement of Science.
Married Diane Joyce Kohn, July 16, 1967. Children: Marcia Ellen, Nicole Elizabeth.