Student, Bryn Mawr College, 1961. Bachelor of Arts in Chemistry cum laude, Radcliffe College, Cambridge, Massachusetts, 1963. Master of Arts in Bacteriology and Biochemistry, Harvard University, Cambridge, Massachusetts, 1965.
Doctor of Philosophy in Microbiology-Molecular Genetics, Harvard University, Cambridge, Massachusetts, 1969. Doctor of Philosophy (honorary), Pierre & Marie Curie University, Paris, 2001. Doctor of Philosophy (honorary), University Copenhagen, 2004.
Milton research fellow, 1969-1970. American Cancer Society research fellow, 1970-1972. Staff biochemist Lawrence Berkeley National Laboratory University California, Berkeley, 1972-1976, member senior staff, 1976, co-director division biology and medicine Laboratory Cell Biology, since 1980, director cell and molecular biology division, 1988-1992, coordinator life science, 1989-1991, associate laboratory director bioscience, 1989, director life science division Lawrence Berkeley National Laboratory, 1992—2002, member faculty department comparative biochemistry, since 1979, associate director biosciences, Lawrence Berkeley National Laboratory, 1995—2002, distinguished scientist, senior advisor to the laboratory director on biology Lawrence Berkeley National Laboratory, since 2002, OBER/Department of Energy Distinguished Scientist Fellow in Life Sciences Lawrence Berkeley National Laboratory, 2005—2010.
Visiting professor Kettering Institute, University Cincinnati Medical Schools, 1986-1988. Distinguished visiting scientist Queensland Institute Medical Research, Brisbane, Australia, 1982. Member council Gordon Research Conference, 1991-1994.
George P. Peacock lecturer pathology University Texas, Dallas, 1992. Dean's lecturer Mount Sinai Medical School, New York City, 1993. Presenter numerous lectures, conductor symposia.
Keynote speaker Gordon Conference on Proteoglycans, 1994, others.
Fellow American Association for the Advancement of Science, Institute of Medicine, American Academy Arts and Sciences, American Philosophical Society. Member National Academy of Sciences, American Society Cell Biology (member council 1989-1991, Women in Cell Biology Career Recognition award 1993, president 1997), International Society Differentiation (board directors 1990-1996, president), American Association Cancer Research (board directors 1999-2001).