Bachelor of Science in Chemistry, Louisiana State University, 1951; Master of Science in Microbiology, Louisiana State University, 1957; Doctor of Philosophy, University Colorado, 1960.
Postdoctoral fellow, National Institute Allergy and Infectious Diseases, U. Oregon, Portland, 1960-1962; microbiologist, Naval Medical Research Institute, Bethesda, Maryland., 1962-1966; research microbiologist, Naval Bioscience Laboratory, Oakland, California, 1966-1967; associate professor medical microbiology and immunology, University of California, Berkeley, 1967-1972; professor, University of California, 1972-1991; professor emeritus, University of California, since 1991. Director Naval Bioscience Laboratory, 1968-1981. Member expert panel on bacteriology World Health Organization, 1972-1991.
Board trustees Alameda (California) Library, 1973-1978. Served to commander Mediterranean Shipping Company United States Naval Reserve, 1952-1955, 62-67. Member American Association Immunologists, American Society Microbiology, International Association Human and Animal Mycology, International Association Microbiol.
Science, Internat.Assn. Aquatic Animal Medicine, Association Military Surgeons.
Married Elizabeth Corbett, April 29, 1955. Children: Sally Ann, Philippa Jane.