Bachelor Pharmacy, London University, 1968. Doctor of Philosophy, University Edinburgh, Scotland, 1971. Doctor of Science, University Edinburgh, Scotland, 1990.
Imperial Chemical Industries research fellow department pharmacology, U. Edinburgh, 1971-1973; Medical Research Council research fellow, U. Edinburgh, 1973-1975; lecturer, U. Edinburgh, 1975-1987; senior lecturer, U. Edinburgh, since 1987; department head, U. Edinburgh, 1995-1998; director studies, U. Edinburgh member department biomedical science, U. Edinburgh, since 1998.
Member British Pharmacol. Society (Sandoz prize), Society for Endocrinology, Society for the Study of Fertility.
Married Valerie Lesley Whitehead, April 19, 1976 (deceased March 1985). Children: Timothy James, Natalie Claire. Married Moira Anderson Scott, August 16, 1990.