Background
He was born in Wiesbaden, Germany.
He was born in Wiesbaden, Germany.
Johns Hopkins University.
He got his early education in private schools in Baltimore, and then spent two years in Baltimore City College (1894-1896). In 1900 he received an Bachelor of Arts degree from Johns Hopkins University. He stayed there to do graduate work, and in 1904 received the degree of Doctor of Philosophy His doctoral thesis as stated in the president"s annual report was entitled "The Larva of Fasciolaria tulipa (var distans)" (or, as published in 1904 in German by Johns Hopkins University Press in Baltimore, Über den kannibalismus bei Fasciolaria tulipa (var distans) und deren larvale excretionsorgane).
In 1901-1902 he was assistant in the United States Bureau of Fisheries and in the North Carolina Geological Survey.
In 1903 he was marine biologist of the Gulf Biological Station and from 1905 to 1907 taught biology at Woods Hole, Massachusetts, during the summers. In 1905, he began teaching zoology at the University of Michigan, becoming in 1908 assistant professor of biology.
In 1916 he moved to Amherst College, where he served as Professor of Biology until 1948 (until 1944 as chairman).