MScAgr, University Helsinki, Finland, 1951. DScAgr, University Helsinki, Finland, 1975.
Researcher, Agricultural Research Center, 1950-1958; research station director, Agricultural Research Center, 1958-1973, 75-80; chief inspector, Ministry Agriculture and Forestry, 1973-1974; professor, U. Helsinki, 1977-1992; director, Seed Testing Station, 1981-1989; director, National Board of Agriculture, 1990-1992; retired, 1992. Lecturer agroecology U. Helsinki, since 1977, professor, since 1981. With Council for Agricultural and Forestry in Finland's Academy, 1980-1985.
Chairman, board directors several schools and municipal bodies. Corporation Anti-Aircraft Artillery, 1944. Member Union of Agricultural Societies (board directors 1976-1991), Finnish Beekeeper's Association (chairman 1986-1991), Kallio (Helsinki) Rotary Club (president 1994-1995).
Married Aino Penttila, 1950 (divorced 1984). Children: Leena, Riitta, Jukka, Matti. Married Pirkko Marttila (deceased 1998).