Bachelor of Science, University Stockholm, 1967. Master of Science, Stanford University, 1968. Doctor of Medicine, University Lund, 1975.
Doctor of Philosophy, University Lund, 1979.
Assistant University Lund, Sweden, 1973-1979. Postdoctoral fellow University Cincinnati, 1980. Assistant professor, associate professor University Lund, 1981-1993, professor, since 1993.
Member Scandinavian Physiological Society, Biophysics Society.
Married Ingegerd Monica Sjöholm, May 26, 1976. Children: Nils, Erik, Eva.