Career
He was a professor at Lund University 1883-1902, later at Uppsala University. He collected plants on a number of journeys to then little-known areas of the world, e.g. Svalbard in 1868, Greenland in 1870 (both times with Nordenskiöld"s expedition) and, in 1873, New Zealand, Australia, Hawaii and California.
He made particularly novel collections and descriptions of bryophytes and to less extent vascular plants, algae and fungi.
His collections are kept at Lund University.