Doctor of Philosophy, State University of New York, 1969.
Postdoc fellow University Kentucky, Lexington, 1969—1971. Assistant investigator Carnegie Institution Washington, Baltimore, 1971—1974. Associate professor Case Wester Reserve University, Cleveland, 1974—1980.
Professor University Cincinnati College Medicine, Cincinnati, since 1980, chairman department cell & cancer biology, 1996—2008.
Married Mary Piper, January 14, 1981. 1 child Elizabeth Piper.