Division on Dynamical Astronomy, Dookie Agricultural College, Australia, 1959. BAgrSci, University Melbourne, 1963. Doctor of Philosophy, University Sydney, 1966.
Master of Science, London University, 1978. Doctor of Science, London University, 1980.
Research fellow Babraham Institute, Cambridge, England, 1967-1969. Research instructor Washington University, St. Louis, 1970. Visiting instructor Northwestern University, Chicago, 1971.
Senior research fellow Oxford (England) University, 1972-1974. Lecturer, senior lecturer Chelsea College, London University, 1975-1985. Reader, professor King's College, London, since 1985.
European Molecular Biology Organization fellow Hadassah Medical School, Jerusalem, 1973. Visiting professor University Guelph, Ontario, Canada, 1988, Robert Gordon University, Aberdeen, Scotland, since 1993, University Pittsburgh, 1993. Monbusho professor Nagoya (Japan) University, 1993.
Member Bio-chemical Society (chairman Harden Conference committee 1986-1993, chairman membrane group 1988-1993).
Married Christine Ann McCabe, March 3, 1962. Children: Dimity Ann, Gray Nicholas, Simon Spencer, Michael Milton.