Bachelor of Science in Zoology, University Illinois, 1963. Doctor of Philosophy in Marine Biology, University Miami, Florida, 1973.
Research associate Academy of Natural Sciences, Philadelphia, 1973-1974. Research scientist Commonwealth Scientific and Industrial Research Organisation division of Fisheries and Oceanography, Cronulla, Australia, 1974-1977. Curator of fishes J.L.B. Smith Institute Ichthyology, Grahamstown, South Africa, 1978—2002, curator emeritus, since 2002.
Consultant fish taxonomist Food and Agriculture Organization Fishery Resources and Environmental Division, Rome, 1980-1996, Namibian Fisheries Department, Windhoek, 1990-1996.
Member American Society Ichthyologists and Herpetologists, Society of Systematic Biologists.
Married Elaine Margaret Lawrence, March 15, 1991. Children: Lydia, Julia, Andrew.