Bachelor of Scientific Agriculture, University Toronto, 1956. Master of Science, University Wisconsin-Madison, 1958. Doctor of Philosophy, University Wisconsin-Madison, 1959.
Doctor of Science (honorary), University Guelph, 1997.
Assistant professor University Guelph, Ontario, Canada, 1960-1963. Research biochemist DuPont deNemours & Company, Wilmington, Delaware, 1963-1967, research supervisor, 1967-1974, associate director, 1974-1979, director life science, 1979-1984. President Bio Technica International, Inc., Cambridge, Massachusetts, 1984-1986.
President, Chief Executive Officer Boyce Thompson Institute, Inc., Ithaca, New York, 1986-1995, president emeritus, since 2000. Deputy chairman Bio Technica International, Inc., 1986-1990, consultant, board directors, 1990-1999. President National Agricultural Biotechnology Council, Ithaca, since 1996.
Member executive committee board agriculture National Research Council, 1982—1988, member commission life science, 1984—1990, board biology, 1984—1990, member committee on biotechnology, 1988—1995, chairman committee, 1993—1994, board science technological international development, 1990—1993, chairman committee on biological control, 1992—1995, chairman committee on biological nitrogen fixation, 1992—1994, chairman committee on natural products, 1996—1997. Member committee genetic experimentation International Council of Scientific Union, 1981—1995. Chairman, founder National Agricultural Biotechnology Council, 1988—1993.
Member science advisory committee United States Department Energy, 1991—1995. Member alternate agriculture research commercial board United States Department of Agriculture, 1992—1996, member and corporation secretary alternate agricultural research commercial corporation, 1996—2000. Member Canada reallocations committee Natural Sciences and Engineering Research Council, 1997—1998.
Member science advisory board Foragen, Guelph, Ontario, Canada, 1999—2006, Agriculture and Agri-Food Canada, since 2004. Board directors BioCap, Canada, 1998—2008.
Member biotechnology Executive Board Cornell Univercity, 1986-1995, advisory county Veterinary College, 1989-1996. Member governor board Cornell Center for Environment, 1991-1995. Member Industrial Biotechnology Association (board directors 1986-1989), Agriculture Research Institute (bd.govs.
1988-1991), American Chemical Society (Executive Committee biological chemistry division 1978-1981, Delaware award 1969), American Society Biological Chemists and Molecular Biologists, American Society Plant Physiology (Executive Committee, treasurer 1974-1977), American Society Agronomy, American Society Microbiology.
Married Jacqueline M. Thayer, December 26, 1954. Children: Steven, Chris, Barbara, Ralph (deceased), Jon.