Career
He received his Bachelor of Arts from Columbia University in 1902 and his Doctor of Medicine degree from the College of Physicians and Surgeons three years later. He published his groundbreaking paper on blood transfusion before World War I. In haemocompatibility tests, which he had started in 1907 he found out that patient antibodies against donor red cells could be harmful but not vice versa. This report led to the use of group O (“zero”) individuals as universal donors.