Bachelor in Biology and Chemistry with honors, McGill University, Montreal, 1924; Master of Arts in Biochemistry, McGill University, Montreal, 1930; Doctor of Philosophy in Experimental Medicine cum laude, McGill University, Montreal, 1932.
Biochemistry research technician, Medical Laboratory, Royal Victoria Hospital, McGill University, Montreal, 1925-1929; demonstrator in physiology, Banting & Best Medical Research Institute, U. Toronto, Ontario, Canada, 1933-1935; researcher on hearing, Physiology Department, McGill University, Montreal, 1936; assistant professor physiology, Medical College, Dalhousie U., Halifax, Nova Scoti, Canada, 1937-1938; teaching and research faculty appointee department physiology, McGill University, 1939-1947; research associate clinical science, medical College University of Illinois, Chicago, 1948-1961; research associate pathology, medical College University of Illinois, 1961-1966. Participant in International Symposium on Gastric Cancer, National Cancer Institute/National Science Foundation, Japan, 1969.
Fellow Royal Society Medicine (England). Member Canada Physiological Society, American Physiological Society, Sigma Xi.