Bachelor of Science, University of Illinois, 1959; Master of Science, University of Illinois, 1962; Doctor of Philosophy in Zoology and Genetics, University of Illinois, 1965.
Research associate mosquito genetics University Illinois, 1965, research associate, 1965-1966, assistant professor mosquito genetics, 1966-1969. Associate professor internal medicine School Medicine University Maryland, Baltimore, professor, director International Health Program, 1979-1982. Director Pakistan Medical Research, Lahore, 1969-1979.
With Florida Medical Entomological Lab, Institute Food and Agriculture Sciences University Florida, Vero Beach, 1982-1999, director Florida Medical Entomological Laboratory, Institute Food and Agriculture Sciences, 1998-1999. Professor emeritus Florida Medical Entomological Laboratory, Institute Food and Agr.Scis., University Florida, since 2000. Consultant World Health Organization, 1965.
Member Genetics Society of America, Entom. Society of America, American Society Tropical Medical and Hygiene, American Genetic Association, American Mosquito Control Association, Sigma Xi.
Married; two children.