Student, St. Olaf College, 1967. Bachelor of Science, University Wisconsin, Eau Claire, 1974. Master of Science, University Montana, 1978.
Doctor of Philosophy, University Chicago, 1982.
Postdoctoral fellow, University of Chicago, 1982-1983; postdoctoral fellow virology, Stanford University, 1983-1986; scientist, Chiron Corporation, Emeryville, California, since 1986.
Served with United States Army, 1968-1970, Vietnam. Member American Association for the Advancement of Science, American Society Microbiology, Society for General Microbiology, American Society for Virology.