AB, Princeton University, 1956; Doctor of Philosophy, Harvard University, 1959.
Assistant professor biophysics, University of Chicago, 1961-1964;
associate professor, University of Chicago, 1964-1969;
professor, department chairman, University of Chicago, 1969-1984;
F.L. Pritzker Distinguished Service professor department molecular genetics and cell biology, University of Chicago, since 1984;
director, Center for Photochemistry and Photobiology, since 1987. Chairman, Board Of Directors Integrated Genomics, Inc., Chicago, since 1997. Consultant virology and rickettsiology study section United States Public Health Service, 1969-1973.
Member science advisory board Sloan-Kettering Institute, 1978-1979. Member nitrogen fixation panel United States Department of Agriculture, 1978-1979. Member panel science advisory United Nations Industrial Development Organization International Center for Genetic Engineering and Biotechnology, 1984-1994, 97-.
Member recombinant DNA advising committee National Institutes of Health, 1991-1995. Adjunct scientist Woods Hole Oceanographic Institution, since 1994.
Trustee Marine Biological Laboratory, Woods Hole, Massachusetts, since 2003. Fellow American Association for the Advancement of Science, American Academy Arts and Sciences (chairman, midwest council, vice president 1993-1999), American Academy Microbiology. Member National Academy of Sciences, International Society Plant Molecular Biology (president 1987-1989).
Married Margot Block, June 23, 1957. Children: Deborah, David.