Bachelor cum laude, Harvard University, 1956. Doctor of Medicine, Harvard University, 1960.
Chief section microbial genetics laboratory of molecular biology National Institute Diabetes and Digestive Disease, National Institutes of Health, Bethesda, Maryland, 1960—1990, biochemist Bethesda, since 1990, Laboratory Molecular Biology, Bethesda, since 1990. Chairman biohazards committee National Institutes of Health, Bethesda, 1972—1975. Surgeon United States Public Health Service, 1960-1990.
Fellow: American Society Experimental Biology, American Academy Microbiology. Member: American Society Microbiology, Genetics Society.
Married Judith Sylvia Perlman, January 30, 1960. Children: Nicholas Ivor, Jacobina Helen.