Bachelor of Science in Chemistry, St. Saint Louis University, 1959; Master of Arts in Medical Scis, Harvard University, 1961; Doctor of Philosophy in Biophysics, Harvard University, 1964.
Postdoctoral fellow, Medical Research Council, National Institutes of Health, Cambridge, England, 1963-1965; postdoctoral fellow, National Academy Sciences-National Research Council, Institut Pasteur, Paris, 1965-1966; professor, chairman molecular biology and biochemistry, University of Wisconsin, Madison, 1966-1981; John G. Searle professor, chairman molecular biology and biochemistry, Medical and Dental Schools, Northwestern University, Chicago, 1981-1990; leader cancer molecular biology program, Cancer Ctr.Northwestern University, Chicago, 1982-1989; professor biochemistry, director Center for Molecular Biology, Wayne State University, Detroit, 1990-1994; interim chair department molecular biology and genetics, Wayne State University, Detroit, 1993-1994; director, professor Center for Molecular Medicine and Genetics, Wayne State University School Medicine, Detroit, 1994-1997; professor internal medicine, Wayne State University School Medicine, Detroit, 1994-1997. Vice chairman Gordon Research Conference Extrachromosomal Elements, 1984, chairman,1986. Honorary research professor Biotechnology Research Center, Chinese Academy Agricultural Sciences, Beijing, 1987-1997.
Member troop committee, treasurer Four Lakes Council Boy Scouts American, Madison, 1973-1977, member People to People Program delegate of microbiologists to China, 1983. Member National Academy of Sciences/National Research Council Committee on Human Health Effects of Subtherapeutic Antibiotic Use in Animal Feeds, 1979-1981. Senior technical advisory recruitment consultant United Nations Development Program in China, 1987.
Member National Institutes of Health (microbial genetics study section 1978-1982, special study section 1971, 73, 74, 75, 78, 79, 82, chairman 1983, 84-86, 88, 89, 90, 93-95, director medical scientist training program 1982-1990, advisory panel National Research Council 1974-1977, chairman 1976-1977, advisory panel for development biology National Science Foundation, 1968-1971, National Science Foundation advisory panel North Atlantic Treaty Organization postdoctoral fellowships in science program 1979), American Society Microbiology, Association Harvard Chemists, American Society Biological Chemists, American Academy Microbiology, New York Academy of Sciences.
Married Rosalie Anne Lowery, June 13, 1959. Children: Jennifer Rose, Robert Harvey, David Matthew.