Bachelor of Science, Brooklyn College, 1963. Doctor of Philosophy, Indiana University, 1968.
Postdoctoral fellow, U. Hawaii, Honolulu, 1968-1970; postdoctoral fellow, Princeton (New Jersey) U., 1970-1971; assistant professor, Boston University, 1971-1977; associate professor, Boston University, 1977-1983; full professor, Boston University, 1983-1996; chairman biology department, Boston University, 1990-1996; dean science, U. Massachusetts, Lowell, since 1996. Member science advisory board Manomet (Massachusetts) Bird Observatory, 1984-1996.
Member American Society Mammalogists (board directors 1979-1985, editorial board 1982-1991), Ecological Society of America (ad hoc editor 1984-1985), Boston Computer Society (International Business Machines Corporation product review editor 1987), Sigma Xi (local chapter president 1981-1982, since 2000).
Married Virginia M. Londy, May 31, 1968. Children: David L., Bonnie.