Bachelor of Science, Brooklyn College, 1962; Doctor of Philosophy, University Washington, 1967.
National Institutes of Health postdoctoral fellow, University of Wisconsin -Madison, 1967-1969;
assistant professor collegiate division biology, University of Wisconsin -Madison, 1969-1970;
assistant professor department biology, University of Chicago, 1970-1975;
associate professor, University of Chicago, 1975-1982;
professor department biology, University of Chicago, 1982-1984;
professor department molecular genetics and cell biology, University of Chicago, since 1984;
National Center for Scientific Research/National Science Foundation fellow, University of Chicago, 1976-1977. Member genetic basis of disease panel National Institutes of Health, 1978-1981, biomedical science panel,1982-1986, chairperson, 1987-1989. Director National Institutes of Health/National Research Service Award Genetics and RegulationTng.
Grant, since 1988, member genome research review committee, since 1991.
Member Genetics Society of America (treasurer 1983-1986), American Society Microbiology, American Association for the Advancement of Science, American Society Cell Biology.