Bachelor of Science, University Lund, 1973. Doctor of Philosophy, University Lund, 1976.
Associate professor medical faculty University Umeå, Sweden, 1980-1986. Associate professor Cancer Research Foundation, Stockholm, 1987-1989. Professor Swedish Natural Science Research Council, 1989-1993.
Professor, chairman development biology University Uppsala, Sweden, since 1993. Department head University Uppsala, since 1994, director microscopy and imaging unit, since 1993.
Member American Association for the Advancement of Science, New York Academy of Sciences, American Association Cancer Research.
Married Susan Bente Pfeifer, June 24, 1982. Children: Alexandra, Sebastian.