Bachelor, University Texas, 1974. Master of Arts, University of California at Los Angeles, 1977. Doctor of Philosophy, Duke University, 1984.
Research assoc Institute Environmental and Evolutional Biology, Los Angeles, 1976-1977. Research assistant Smithsonian Institution, Washington, 1977-1979. Postdoctoral fellow Smithsonian Marine Station, Fort Pierce, Florida, 1984-1985.
Professor Millsaps College, Jackson, Mississippi, since 1985. Consultant curriculum revision Rice University, Houston, 1991-1994, curriculum materials Abt Associates, Cambridge, Massachusetts, since 1996.
Member American Association for the Advancement of Science, Society Integrative and Comparative Biology, Sigma Xi.