Bachelor of Science, University Western Australia, 1964. Doctor of Philosophy, Monash University, Melbourne, Australia, 1968.
Lecturer Flinders University, Australia, 1968-1969. Research fellow Carnegie-Mellon University, Pittsburgh, 1969-1972. Research scientist Roswell Park Cancer Institute, Buffalo, since 1972.
Professor biophysics State University of New York, since 1984, chair molecular cell biophysics department, 1988—2004, senior research scientist cell biological department, since 2005. Member high voltage electron microscope advisory committee National Institutes of Health, Albany, New York, 1981-1984, member biophysics chemistry study section, 1986-1990.
Fellow American Physical Society. Member Biophysics Society, Microscopy Society American (board directors 1984-1987), American Society for Cell Biology.
Married Hilda Liang, December 18, 1971. Children: Jennifer S., Peter S.