Career
He discovered double fertilization in plants in 1898. 1874 — enters the Medical Surgical Academy in Saint St. Petersburg, works on chemistry in the laboratory of A. Borodin
1878 — moves to the Moscow University, obtains Candidate degree in 1881 in Chemistry. Under the influence of K. Timiryazev and V. Zinger starts to study Botany.
Receives a position of a laboratory assistant at the chair of Plant Physiology and later (1885) in the Petrovskaya Agricultural Academy.
1894 — is invited to work at the chair of Systematics and Morphology of the Kiev University. During 1894-1914 works as a director of the Botanical Garden of Kiev University
1918-1923—professor of Tbilisi University (Georgia)
1923—founds the Timiryazev Biological Institute in Moscow.
Heads it till 1929.
