Bachelor of Science, Haverford (Pennsylvania) College, 1973; Doctor of Philosophy, Yale University, 1979; Doctor of Medicine, Yale University, 1980.
Instructor in medicine, Harvard Medical School, Boston, 1985-1989; assistant professor cell molecular physiology, Harvard Medical School, Boston, 1989-1993; assistant professor cell biology, Harvard Medical School, Boston, 1993-1995; associate professor cell biology (medicine), Harvard Medical School, Boston, since 1995; assistant physician, Beth Israel Hospital, Boston, since 1992.
Fellow American Association for the Advancement of Science (medical science section).
Married Roberta S. Isberg, May 29, 1983. Children: Rebecca Sylvia, Harry Joseph.