AB, Princeton University, 1954; Doctor of Medicine, Harvard Medical School, 1959; Doctorate (honorary), University Tampere, 1992.
Intern, Bellevue Hospital, New York City, 1959-1960; research scientist, National Institute of Mental Health, Bethesda, Maryland., 1960-1962; research specialist, National Institute of Mental Health, Bethesda, Maryland., 1964-1978; lecturer, Georgetown University Department Physiology, Washington, 1970-1984; chief Laboratory Neuroscis., National Institutes of Health National Institute Aging., Bethesda, since 1978. Member of advisory board Brain Dysfunction, Italy, since 1984. Member of advisory panel Fogerty Schools Residence Program, Bethesda, 1985-1989.
Fellow American College Neurophychopharmacology, Royal Society Medicine, Gerontological Society American. Member Alzheimers Disease Association (member medical advisory board 1985-1989).
Married Judith Helen Livant, June 25, 1961. Children; Erik Alexander, Stuart Daniel.