University of Alaska (Bachelor of Arts). University of California Boalt Hall School of Law (Bachelor of Laws).
National Science Foundation fellow University Microbiology Institute, Copenhagen, 1963-1964, Hammersmith Hospital, London, 1964-1965. Research associate Tufts Medical School, Boston, 1965-1966. Assistant research professor State University of New York-Buffalo, 1966-1970.
Associate professor University Michigan Medical School, Ann Arbor, 1970-1978, professor microbiology, since 1978. Fogarty International fellow Imperial Cancer Research Fund, London, 1976.
Member American Microbiology Society, American Association for the Advancement of Science.
Married Alexandra Lee Winer, August 21, 1960. Children: Eric, Michael.