AB, University of California, Berkeley, 1974; Doctor of Philosophy, University of California, Riverside, 1981.
Postdoctoral fellow, Albert Einstein College Medicine, New York City, 1981-1982; postdoctoral fellow, Fred Hutchinson Cancer Research Center, Seattle, 1982-1986; assistant professor cellular biology, Vanderbilt University School Medicine, Nashville, 1986-1993; associate professor, Vanderbilt University School Medicine, Nashville, since 1993. Consultant Upstate Biotechnology Inc., Lake Placid, New York, since 1993.
Married Camille Marie O'Kara, December 22, 1981 (divorced January 1989). 1 child, Trevor David.